1. Department of Biology, Faculty of Science, Prince of Songkla University, Songkhla, Hat Yai, Thailand
2. Department of Science, Wangbama Central School, Ministry of Education, Thimphu, Bhutan
3. Conservation Ecology Program, School of Bioresources and Technology, King Mongkut’s University of Technology, Thonburi, Bangkok, Thailand
4. Tsirang Forest Division,Tsirang, Department of Forest and Park Services, Ministry of Agriculture and Forests, Bhutan
5. Institute of Botany and Landscape Ecology, University of Greifswald, Greifswald, Germany
6. Nature Conservation Division, Department of Forest and Park Services, Ministry of Agriculture and Forests, Thimphu, Bhutan
7. Science Research Center, Hosei University, Fujimi 2-17-1 Chiyoda-ku, Tokyo 102-8160, Japan