1. College of Sciences of Tunis, Tunis El Manar University, Tunis, Tunisia
2. Laboratory of Aromatic and Medicinal Plants (LPAM), Biotechnology Center in Borj Cedria Technopole, Hammam-Lif, Tunisia
3. Espace d’Appui a` la Recherche et Transfert Technologique (ARTT), Centre de Biotechnologie de Borj Cedria, Hammam-Lif, Tunisia
4. Laboratory of Bioactive Substances (LSBA), Biotechnology Center of Borj-Cedria, Hammam-lif, Tunisia
5. Department of Biology, College of Sciences, Taif University, Taif, Saudi Arabia