1. Department of Genetics, College of Life Science, Northeast Forestry University, Harbin, PR China;
2. Jilin Province Product Quality Supervision and Inspection Institute, Changchun, PR China;
3. Institute of Plateau Ecology, Tibet Agriculture and Animal Husbandry College, Linzhi, PR China;
4. Institute of Rural Energy and Ecology, Jilin Academy of Agricultural Science, Changchun, PR China;
5. Medicinal Plants Research Center, Tibet Agriculture and Animal Husbandry University, Nyingchi, PR China