1. a Veterinary Sciences Division , Department of Agriculture and Rural Development for Northern Ireland , Stormont , Belfast , BT4 3SD , UK
2. b Department of Veterinary Science , Queen's University of Belfast , Stormont , Belfast , BT4 3SD , UK
3. c Dipartimento di Scienze Biopatologiche ed Igiene delle Produzioni Animali e Alimentari , Università degli Studi di Perugia , Via S. Costanzo 4, 06126 , Perugia , Italy
4. d California Animal Health and Food Safety Laboratory System , Fresno Branch, University of California , Davis, 2789 South Orange Avenue, Fresno , CA , 93725 , USA
5. e Department of Wildlife Diseases , National Veterinary Institute , 75189 , Uppsala , Sweden