1. Washington Cooperative Fish and Wildlife Research Unit, School of Aquatic and Fishery SciencesUniversity of WashingtonBox 355020SeattleWashington98195USA
2. U.S. Geological Survey, Washington Cooperative Fish and Wildlife Research Unit, School of Aquatic and Fishery SciencesUniversity of WashingtonBox 355020SeattleWashington98195USA
3. Lower Elwha Klallam TribeNatural Resource Department51 Hatchery RoadPort AngelesWashington98363USA
4. School of Aquatic and Fishery SciencesUniversity of WashingtonBox 355020SeattleWashington98195USA