1. School of Biology and Center of Excellence in Phylogeny of Living Organisms, College of Sciences, University of Tehran, Tehran, Iran
2. Department of Biology, Faculty of Sciences, Gonbad Kavous University, Golestan, Iran
3. Department of Biology, Faculty of Sciences, Semnan University, Semnan, Iran
4. Insect Taxonomy Research Department, Iranian Research Institute of Plant Protection, Tehran, Iran
5. Institute of Ecology, Evolution and Diversity, Goethe University, Frankfurt/M, Germany
6. Senckenberg Biodiversity and Climate Research Centre, Senckenberg Gesellschaft für Naturforschung, Frankfurt/M, Germany