1. Centre for Ecological Genomics and Wildlife Conservation, Department of Zoology, University of Johannesburg, Auckland Park, 2006, South Africa
2. Department of Biodiversity and Conservation Biology, University of the Western Cape, Bellville, Cape Town, South Africa
3. Department of Forestry, Fisheries and the Environment, Oceans & Coasts Branch, Oceans & Coasts Research, Private Bag X4390, Cape Town, 8001, Western Cape, South Africa
4. Department of Biological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, Cape Town, 7701, South Africa
5. Department of Forestry, Fisheries and the Environment, Fisheries Branch, Fisheries Research, Private Bag X2, Roggebaai, Cape Town, 8012, Western Cape, South Africa