1. Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand
2. Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration, PO Box 582, No. 86, Norodom Blvd, Phnom Penh, Cambodia
3. Wonders of the Mekong Project, PO Box 582, c/o IFReDI, No. 86, Norodom Blvd, Phnom Penh, Cambodia
4. Department of Environmental Biology, Chubu University, Kasugai, 487-8501, Japan
5. Academy of Science, The Royal Society of Thailand, Bangkok, 10300, Thailand