1. Molecular Invertebrate Systematics and Ecology Laboratory, Graduate School of Engineering and Science, University of the Ryukyus, Nishihara, 903-0213, Okinawa, Japan
2. Dipartimento di Scienze della Vita e dell’Ambiente (DiSVA), Università Politecnica delle Marche, Via Brecce Bianche sn, Ancona, 60131, Italy
3. Department of Zoology, National Museum of Nature and Science, Tsukuba, 305-0005, Ibaraki, Japan
4. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, 113-0003, Japan
5. Department of Biochemistry, Genetics and Immunology, University of Vigo, Vigo, Spain
6. Tropical Biosphere Research Center, University of the Ryukyus, Nishihara, 903-0213, Okinawa, Japan