1. Department of Biodiversity, Ecology and Evolution, Faculty of Biology, Universidad Complutense de Madrid, Madrid, Spain
2. Departamento de Ecoloxía e Bioloxía Animal, Universidade de Vigo, Vigo, E-36310, Spain
3. Institute of Evolutionary Biology (CSIC-Universitat, Pompeu Fabra), Passeig Marítim de la Barceloneta 37-,49, Barcelona, 08003, Spain