1. Key Laboratory of Orogenic Belts and Crustal Evolution, School of Earth and Space Sciences, Peking University, 5 Yiheyuan Road, Beijing, 100871
2. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China
3. CAS Center for Excellence in Life and Paleoenvironment, Beijing, China
4. National Museum of Natural History, Bulgarian Academy of Sciences, 1, Blvd. Tzar Osvoboditel, 1000 Sofia, Bulgaria,