1. Department of Earth Sciences, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario, K1S 5B6, Canada,
2. Museum für Naturkunde, Leibniz Institute on Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany,
3. Department of Biology, Humboldt University, Invalidenstrasse 42, 10115 Berlin, Germany
4. Department of Biology, University of Toronto at Mississauga, 3359 Mississauga Road, Mississauga, Ontario, L5L 1C6, Canada,
5. International Center of Future Science, Dinosaur Evolution Research Center, Jilin University, Changchun 130061, China