1. a UPR 2147 du CNRS, 44 rue de l’Amiral Mouchez , 75014 , Paris , France
2. b Institute of Human Origins, Arizona State University , Box 874101, Tempe , Arizona , 85282 , U.S.A.
3. c Department of Anthropology and Center for the Advanced Study of Hominid Paleobiology , George Washington University , Washington , D.C. , 20052 , U.S.A.