1. Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, U.S.A.,
2. Geosciences Program, Queensland Museum, Brisbane, Queensland 4011, Australia
3. Evolutionary Studies Institute, School of Geosciences, University of the Witwatersrand, Private Bag 3, WITS 2050, Johannesburg, South Africa, , ,
4. Natural History Museum, London SW7 5BD, United Kingdom,