1. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science , Invalidenstrasse 43, 10115 Berlin, Germany
2. Department of Mammalogy, American Museum of Natural History , Central Park West at 79th Street, New York, New York 10024, U.S.A.
3. Institute of Human Origins, School of Human Evolution and Social Change, Arizona State University , Tempe, Arizona 85287, U.S.A.,