1. Iwate University, Kurume Fukuoka, Japan
2. Maize and Cash Crop Research Center, National Agricultural and Forestry Research Institute, Xaythany, Vientiane Capital, Lao Democratic People's Republic
3. Priority Organization for Innovation and Excellence, Kumamoto University, Kumamoto, Japan
4. Present address: Faculty of Advanced Science and Technology, Kumamoto University, Kumamoto, Japan