1. College of Natural Resources and Environment, Northwest A&F University, Yangling, Shaanxi, China
2. College of Life science, Tibet University, Tibet, China
3. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences, Fudan University, Shanghai, China
4. Institute of Soil and Water Conservation, Chinese Academy of Sciences, Ministry of Water Resources, Yangling, Shaanxi, China