1. Department of EcologyMontana State UniversityMontana Department of Fish, Wildlife and ParksBozemanMontana59717USA
2. Wildlife Conservation Society301 North Willson Avenue. 59715BozemanMontanaUSA
3. Montana Department of Fish, Wildlife and Parks39 Centerville Road. 59644TownsendMontanaUSA
4. Department of EcologyMontana State University59717BozemanMontanaUSA
5. U.S. Geological Survey, Montana Cooperative Fishery Research UnitMontana State University59717BozemanMontanaUSA