1. Centre for Research and Education on Biological Evolution and Environment, School of Earth Sciences and Engineering, Nanjing University, Nanjing, China
2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China
3. Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, China
4. Department of Earth Sciences, Natural History Museum of London, London, UK