1. Kunming geological Survey of Natural Resources Center, Kunming, China
2. Canadian Museum of Nature, Ottawa, Canada
3. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, P.R. China
4. CAS Center for Excellence in Life and Paleoenvironment, Beijing, P.R. China
5. College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, P.R. China