1. Department of Environmental and Evolutionary Biology, University of Glasgow
UK E-mail: j.hughes@bio.gla.ac.uk (J.H.)
2. Allan Wilson Centre for Molecular Ecology and Evolution, Department of Zoology, University of Otago
P.O. Box 56, Dunedin, New Zealand
3. Illinois Natural History Survey
Champaign, IL 61820, USA
4. Museum of New Zealand Te Papa Tongarewa
Wellington, New Zealand