1. Department of Microbiology, College of Medicine and Medical Research Institute, Chungbuk National University, Cheongju, Republic of Korea
2. Department of Biochemistry, College of Medicine and Medical Research Institute, Chungbuk National University, Cheongju, Republic of Korea
3. Center for Study of Emerging and Re-emerging Viruses, Korea Virus Research Institute, Institute for Basic Science (IBS), Daejeon, Republic of Korea