1. Ashoro Research Forest, Kyushu University, Ashoro, Japan
2. Graduate School of Agricultural Science, Kobe University, Kobe, Japan
3. Miyazaki University Forests, University of Miyazaki, Tano, Japan
4. Department of Environmental Sciences and Technology, Kagoshima University, Kagoshima, Japan
5. Graduate School of Agriculture, Ehime University, Matusyama, Japan