1. Department of Geology and Palaeontology, Muséum d’Histoire Naturelle, CP6434, 1211 Geneva 6, Switzerland;
2. Department of Biology, Faculty of Science, Mahasarakham University, Khamriang, Kantarawichai District, Maha Sarakham 44150, Thailand;
3. Palaeontological Research and Education Centre, Mahasarakham University, Khamriang, Kantarawichai District, Maha Sarakham 44150, Thailand;
4. Palaeobiosphere Evolution, OD Earth and History of Life, Royal Belgian Institute of Natural Sciences, Rue Vautier 29, 1000 Brussels, Belgium