1. Shaanxi Key Laboratory of Early Life and Environments, State Key Laboratory of Continental Dynamics, and Department of Geology, Northwest University, Xi’an, 710069, PR China
2. Department of Zoology, University of Cambridge, Downing Street, Cambridge, CB2 3EJ, UK
3. Centre for Ecology and Conservation, University of Exeter, Penryn Campus, Treliever Road Penryn, Cornwall, TR10 9FE, UK
4. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, PR China