1. Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 92 Box, No. 1 Beichen West Road, Chaoyang District, Beijing, 100101, PR China
2. University of the Chinese Academy of Sciences, Yuquan Road, Shijingshan, Beijing, 100039, PR China
3. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China
4. College of Plant Protection, Agricultural University of Hebei, Baoding, 071001, PR China