1. School of Biological Sciences and Swire Institute of Marine Science, The University of Hong Kong, Kadoorie Biological Sciences Building, Pokfulam Road, Hong Kong SAR, China
2. Camborne School of Mines, Environment and Sustainability Institute, University of Exeter, Penryn Campus, Penryn, Cornwall, TR10 9FE, UK
3. Department of Geological Sciences, University of Florida, 241 Williamson Hall, PO Box 112120, Gainesville, FL 32611-2120, USA
4. Florida Museum of Natural History, 1659 Museum Road, Dickinson Hall, Gainesville, FL 32611, USA