1. Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun, Jilin, PR China
2. University of Chinese Academy of Sciences, Beijing, PR China
3. Jilin Provincial Joint Key Laboratory of Changbai Mountain Wetland and Ecology, Changchun, Jilin, PR China
4. U.S. Geological Survey, Wetlands and Aquatic Research Center, Lafayette, LA, USA