1. Department of Molecular Medicine and Haematology, Faculty of Health Sciences and the Evolutionary Studies Institute, Faculty of Science, University of the Witwatersrand, Johannesburg, 2193, South Africa
2. Agricultural Research Council, Biotechnology Platform, Pretoria 0040, South Africa
3. Department of Biological Sciences, Graduate School of Science, University of Tokyo, Tokyo 113-0033, Japan
4. Department of Biodiversity and Conservation Biology, Faculty of Natural Sciences, University of the Western Cape, Cape Town, 7530, South Africa
5. Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, AZ 85721, USA
6. Department of Biology, University of Western Ontario, London, ON, N6A 5B7, Canada