1. Evolutionary Studies Institute (ESI), University of the Witwatersrand, Johannesburg, South Africa
2. Unit for Environmental Sciences and Management, Faculty of Natural and Agricultural Science, North-West University, Potchefstroom, South Africa
3. Applied Behavioural Ecology and Ecosystem Research Unit, School of Environmental Sciences, University of South Africa, Johannesburg, South Africa
4. Elephants Alive, Hoedspruit, South Africa
5. School of Animal, Plant, and Environmental Sciences, University of the Witwatersrand, Johannesburg, South Africa