1. Institute of Integrative Biology, University of Liverpool, Liverpool, UK;
2. Computational Biology Facility, Technology Directorate, University of Liverpool, Liverpool, UK;
3. Department of Mechanical, Materials and Aerospace Engineering, School of Engineering, University of Liverpool, Liverpool, UK;
4. Liverpool Heart and Chest Hospital, Liverpool, UK;
5. Department of Veterinary Pathology and Public Health, Institute of Veterinary Science, University of Liverpool, Liverpool, UK