1. School of Marine Science and Technology, Harbin Institute of Technology, Weihai, People's Republic of China
2. School of Municipal and Environmental Engineering, Harbin Institute of Technology, Harbin, People's Republic of China
3. Key Laboratory of Marine Genetic Resources-State Key Laboratory Breeding Base, Third Institute of Oceanography of State Oceanic Administration, Fujian Provincial Key Laboratory of Marine Genetic Resources, Xiamen, People's Republic of China