1. Department of Biological Sciences, School of Science, The University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan
2. International Center for Island Studies Amami Station, Kagoshima University, Naze-Minatomachi, Amami, Kagoshima 894-0026, Japan
3. Coastal Branch of Natural History Museum and Institute, Chiba, Katsuura, Chiba 299-5242, Japan
4. Okinawa Churaumi Aquarium, Motobu, Okinawa 905-0206, Japan