1. National Institute of Polar Research, 10-3 Midori-cho, Tachikawa, Tokyo 190-8518, Japan
2. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo, Tokyo 113-0033, Japan
3. Department of Geology and Paleontology, National Museum of Nature and Science, Tsukuba, Ibaraki 305-0005, Japan
4. Tateyama Marine Laboratory, Marine and Coastal Research Center, Ochanomizu University, 11 Kouyatsu, Tateyama, Chiba 294-0301, Japan