1. Hunan Fisheries Science Institute, 728 Shuanghe Road Changsha 410153, P. R. China
2. Collaborative Innovation Center for Efficient and Health Production of Fisheries in Hunan Province, 3150 Dongting Avenue, Changde 415000, P. R. China
3. Kunming Natural History of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, P. R. China
4. Kunming College of Life Science, University of Chinese Academy of Sciences, Kunming 650204, P. R. China