1. Department of Biology, Keio University, 4-1-1 Hiyoshi, Yokohama 223-8521, Japan
2. Department of Biosciences and Bioinformatics, Faculty of Science and Technology, Keio University, 3-14-1 Hiyoshi, Yokohama 223-8522, Japan
3. Faculty of Environment and Information Studies, Keio University, 5322 Endo, Fujisawa, Kanagawa 252-0882, Japan
4. National Institute of Advanced Industrial Science and Technology, 2-17-2-1 Tsukisamu-Higashi, Sapporo 062-0052, Japan
5. British Antarctic Survey, High Cross, Madingley Road, Cambridge CB3 0ET, UK
6. National Institute of Genetics, Mishima, Shizuoka 411-8540, Japan
7. National Institute of Polar Research, 10-3 Midori-cho, Tachikawa, Tokyo 190-8518, Japan