1. Department of Biological Sciences, University of Cape Town, Private Bag X3, Rondebosch 7701, South Africa
2. Marine Research Institute, University of Cape Town, Private Bag X3, Rondebosch 7701, South Africa
3. Oceanographic Research Institute, PO Box 10712, Marine Parade, Durban 4056, South Africa
4. Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X17, Bellville 7535, South Africa
5. Department of Earth Sciences, Stellenbosch University, Private Bag X1, Matieland 7602, South Africa
6. Department of Oceanography, University of Cape Town, Private Bag X3, Rondebosch 7701, South Africa