1. a Elwandle Node, South African Environmental Observation Network (SAEON) , Private Bag 1015, Grahamstown , 6140 , South Africa
2. b Department of Ichthyology and Fisheries Science , Rhodes University , PO Box 94, Grahamstown , 6140 , South Africa
3. c Marine and Coastal Management, Department of Environmental Affairs , Private Bag X2, Rogge Bay , 8012 , South Africa
4. d Marine Research Institute, Department of Zoology , University of Cape Town , Private Bag X3, Rondebosch , 7700 , South Africa